When some marsupials developed anteriorly procumbent lower incisors
There are two clades of marsupials in the LRT.
Those with and those without anteriorly oriented = extremely procumbent lower incisors. Kangaroos (Fig 3) arose from the former (with procumbent incisors). Human ancestors arose from the latter (with typical primitive vertical incisors).
The point at which this split arose is documented in the following two sister clades (Figs 1, 2) with Acristatherium and Asioryctes (the unnumbered long rostrum specimen) as outgroup taxa.
Figure 1. Acristatherium and Asioryctes are basal to Proargyrolagus, Tarispes, Caenolestes, Rhyncholestes. Note the anteriorly-projected lower incisors.
” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/rhyncholestes_caenolestes_tarsipes_skull588.jpg?w=116″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/rhyncholestes_caenolestes_tarsipes_skull588.jpg?w=397″ class=”size-full wp-image-90412″ src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/rhyncholestes_caenolestes_tarsipes_skull588.jpg” alt=”Figure 1. Acristatherium and Asioryctes are basal to Proargyrolagus, Tarispes, Caenolestes, Rhyncholestes. Note the anteriorly-projected lower incisors. ” width=”584″ height=”1508″ srcset=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/rhyncholestes_caenolestes_tarsipes_skull588.jpg?w=584&h=1508 584w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/rhyncholestes_caenolestes_tarsipes_skull588.jpg?w=58&h=150 58w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/rhyncholestes_caenolestes_tarsipes_skull588.jpg?w=116&h=300 116w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/rhyncholestes_caenolestes_tarsipes_skull588.jpg 588w” sizes=”(max-width: 584px) 100vw, 584px” />
Figure 1. Acristatherium and Asioryctes are basal to Proargyrolagus, Tarispes, Caenolestes, Rhyncholestes. Note the anteriorly-projected lower incisors.
Other Acristatherium descendants in the LRT
include those in figure 2.
Figure 2. Here Acristatherium and Asioryctes are basal to Palaeothentes, Zalambdalestes, and tiny Zanycteris (not an archaic primate).
” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/zanycteris.zalambdalestes588.jpg?w=160″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/zanycteris.zalambdalestes588.jpg?w=547″ class=”size-full wp-image-90414″ src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/zanycteris.zalambdalestes588.jpg” alt=”Figure 2. Here Acristatherium and Asioryctes are basal to Palaeothentes, Zalambdalestes, and tiny Zanycteris (not an archaic primate). ” width=”584″ height=”1093″ srcset=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/zanycteris.zalambdalestes588.jpg?w=584&h=1093 584w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/zanycteris.zalambdalestes588.jpg?w=80&h=150 80w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/zanycteris.zalambdalestes588.jpg?w=160&h=300 160w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/zanycteris.zalambdalestes588.jpg 588w” sizes=”(max-width: 584px) 100vw, 584px” />
Figure 2. Here Acristatherium and Asioryctes are basal to Palaeothentes, Zalambdalestes, and tiny Zanycteris (not an archaic primate).
… and those in figure 3.
Figure 2. At left, the extant kangaroo, Macropus, and its ancestor, Lagostropheus. At right, the extinct short-faced kangaroo, Procoptodon, and its relative in the LRT, Dendrolagus. Skulls scaled to a common lenglth, making the short-faced kangaroo actually a deep-faced kangaroo. Subtle differences lump and separate these taxa in the LRT.
” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/05/procoptodon-dendrolagus588-1.gif?w=300″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/05/procoptodon-dendrolagus588-1.gif?w=584″ class=”size-full wp-image-86221″ src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/05/procoptodon-dendrolagus588-1.gif” alt=”Figure 2. At left, the extant kangaroo, Macropus, and its ancestor, Lagostropheus. At right, the extinct short-faced kangaroo, Procoptodon, and its relative in the LRT, Dendrolagus. Skulls scaled to a common lenglth, making the short-faced kangaroo actually a deep-faced kangaroo. Subtle differences lump and separate these taxa in the LRT.” width=”584″ height=”511″ srcset=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/05/procoptodon-dendrolagus588-1.gif?w=584&h=511 584w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/05/procoptodon-dendrolagus588-1.gif?w=150&h=131 150w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/05/procoptodon-dendrolagus588-1.gif?w=300&h=263 300w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/05/procoptodon-dendrolagus588-1.gif 588w” sizes=”(max-width: 584px) 100vw, 584px” />
Figure 3. At left, the extant kangaroo, Macropus, and its ancestor, Lagostropheus. At right, the extinct short-faced kangaroo, Procoptodon, and its relative in the LRT, Dendrolagus. Skulls scaled to a common lenglth, making the short-faced kangaroo actually a deep-faced kangaroo. Subtle differences lump and separate these taxa in the LRT.
… and those in figure 4.
Figure 4. The cuscus (genus: Phalanger orientalis) nests with Petaurus and Thylacoleo in the LRT. Note the large lower incisors.
” data-medium-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/phalanger_orientalis588.jpg?w=300″ data-large-file=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/phalanger_orientalis588.jpg?w=584″ class=”wp-image-90477 size-full” src=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/phalanger_orientalis588.jpg” alt=”Figure 4. The cuscus (genus: Phalanger orientalis) nests with Petaurus and Thylacoleo in the LRT. Note the large lower incisors.” width=”584″ height=”410″ srcset=”https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/phalanger_orientalis588.jpg?w=584&h=410 584w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/phalanger_orientalis588.jpg?w=150&h=105 150w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/phalanger_orientalis588.jpg?w=300&h=211 300w, https://pterosaurheresies.wordpress.com/wp-content/uploads/2024/12/phalanger_orientalis588.jpg 588w” sizes=”(max-width: 584px) 100vw, 584px” />
Figure 4. The cuscus (genus: Phalanger orientalis) nests with Petaurus and Thylacoleo in the LRT. Note the large lower incisors.
… and even the one in figure 5.
Figure 5. The marsupial lion, Thylacoleo, also has large incisors, but here the lowers are rotated to become puncturing teeth.
This appears to be a novel hypothesis of interrelationships.
If not, please provide a citation so I can promote it here.
Source: https://pterosaurheresies.wordpress.com/2024/12/16/when-some-marsupials-developed-anteriorly-procumbent-lower-incisors/
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Gee, I’ve always wanted an answer to this question because I lay awake at night trying solve the riddle.